Colonising species have been widely used to search mechanisms of adaptation and speciation due to their merit as biological mateerial. This project aims to investigate these mechanisms using two colonising species of Drosophila, a genus that has demonstrated its value in evolutionary research. The present work will be carried out by two widely eaxperienced teams in ecolutionary biology that join efforts in a coorinated venture. Group 1 has contributed in a significant way to the knowoledge of selecction mechanisms in natural and experimental populations, to the desciphering of generic architecture of reproductive isolation in species hybrids, and to phylogenetic relationships,using D. buzzatii, a South American species that has colonised in historial times the Mediterranean basin. Group 2 has thoroughy studied adpatation at different levels (chromosomal, morphological and molecular) in D. subobscura populations. This group has studied the recent colonisation of different areas of the Americas by this European species, showing that D.subobscura has developped rapidly in colonised areas a latitudinal clinal variation similar to that found in its ancient European range of distribution.This constitutes a unique grand experiment in nature, showing the strength of natural selection, whose meaning for evolutionary theory is going to be investigated more thotough by both groups in this project. Morphological, chromosomal land molecular clinal cariation will be investigated in natural and experimental populations, focusing in the putative role of temperature in shaping this variation (objetive 1). Significantly, observed changes in chromosomal variation in recent years coud be an indicator of global warming if this thermal relationship could be asessed. Several aspects of founder events of founder events such as lethal persistence (objetive 2), transposable element distribution (objetive3), and molecular variation (objetive6) will be investigated, as well.
|Effective start/end date||19/12/00 → 19/12/03|
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